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The 1S hyperfine splitting in hydrogen is measured to an impressive ppt precision and will soon be measured to ppm precision in muonic hydrogen. The latter measurement will rely on theoretical predictions, which are limited by knowledge of the proton polarizability effect Δpol. Data-driven evaluations of Δpol have long been in significant tension with baryon chiral perturbation theory. Here we present improved results for Δpol driven by new spin structure data, reducing the long-standing tension between theory and experiment and halving the dominating uncertainty in hyperfine splitting calculations. 

Abstract Pear is a major fruit tree crop distributed worldwide, yet its breeding is a very time-consuming process. To facilitate molecular breeding and gene identification, here we have performed genome-wide association studies (GWAS) on eleven fruit traits. We identify 37 loci associated with eight fruit quality traits and five loci associated with three fruit phenological traits. Scans for selective sweeps indicate that traits including fruit stone cell content, organic acid and sugar contents might have been under continuous selection during breeding improvement. One candidate gene, PbrSTONE , identified in GWAS, has been functionally verified to be involved in the regulation of stone cell formation, one of the most important fruit quality traits in pear. Our study provides insights into the complex fruit related biology and identifies genes controlling important traits in pear through GWAS, which extends the genetic resources and basis for facilitating molecular breeding in perennial trees.